B.S. California Inst. of Technology, Independ. Studies, Biochemistry (1973) Ph.D.: Yale University, Cell and Developmental Biology (1979) Presidential Young Investigator, National Science Foundation (1984) MacArthur Prize, MacArthur Foundation (1992-1997) Member, National Academy of Sciences (1993-) Fellow, American Academy of Arts and Sciences (1994) George Morel Memorial Fellowship, INRA, France (1998) Distinguished Alumni Award, California Institute of Technology (1998)